Nathan Fillion Recalls Working With Meghan Markle as a Princess on ‘Castle’ (Exclusive)

Nathan Fillion remembers once Meghan Markle was a princess.

The star of the upcoming completely new crime drama The Rookie spoke to ET’s Leanne Aguilera on Tuesday at the ABC upfront event in completely new York City, where he recalled her role on the beloved (nevertheless sadly canceled) crime series Castle. In 2012, Markle appeared in a one-off, fairy tale-themed episode that will saw the likes of Snow White along with Red Riding Hood getting murdered.

As for the future royal family member? She played Sleeping Beauty. Fillion fills inside the spoiler-filled details.

“I remember the entire episode along with I remember she was one of our fairy-tale princesses,” he told ET. “I forgot that will she was the murderer along with I am shocked! I am shocked!”

Fillion then feigned surprise at Markle’s much-anticipated royal wedding coming up This specific Saturday.

“What a wonderful surprise!” he joked. “How lovely for her! Congratulations.”

Markle has popped up in a few surprising places in her acting career outside of her starring role on Suits along with as a heartless fairy-tale princess murderer. She made a brief appearance as a FedEx delivery driver in Horrible Bosses, she was a suitcase design on Deal or No Deal, she played “hot girl” alongside Ashton Kutcher inside the 2005 romantic comedy A Lot Like Love along with appeared as the inquisitive FBI Agent Jessup in an episode of Fringe.

Fillion’s completely new crime drama, The Rookie, airs This specific fall on ABC.

For a closer look at some of Markle’s surprising celebrity connections, watch the video below.

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